1. South China Sea Fisheries Research Institute, Chinese Academy of Fishery Sciences, Guangzhou 510300, China
2. Key Laboratory of Open-Sea Fishery Development, Ministry of Agriculture and Rural Affairs, Guangzhou 510300, China
3. Research and Development Center for Tropical Aquatic Products, South China Sea Fisheries Research Institute, Chinese Academy of Fishery Sciences, Sanya 572018, China
4. Sanya Tropical Fisheries Research Institute, Sanya 572018, China
5. School of Marine Engineering Equipment, Zhejiang Ocean University, Zhoushan 316022, China