1. Department of Global Agricultural Sciences, Graduate School of Agricultural and Life Sciences, The University of Tokyo, Tokyo 113-8657, Japan
2. Department of Agronomy, Faculty of Agriculture, University of Jaffna, Jaffna 40000, Sri Lanka
3. The University of Tokyo Hokkaido Forest, Graduate School of Agricultural and Life Sciences, The University of Tokyo, Furano 079-1563, Hokkaido, Japan