1. School of Agriculture and Food Sustainability, The University of Queensland, Gatton 4343, Australia
2. Unité des Virus Émergents (UVE: Aix-Marseille Univ, Università di Corsica, IRD 190, Inserm 1207, IRBA), 13005 Marseille, France
3. School of the Environment, The University of Queensland, Brisbane 4072, Australia