1. College of Horticulture and Landscape Architecture, Southwest University, Chongqing 400715, China
2. Key Laboratory of Agricultural Biosafety and Green Production of Upper Yangtze River, Ministry of Education, Chongqing 400715, China
3. Laboratoire Universitaire de Biodiversité et Écologie Microbienne, University Brest, INRAE, F-29280 Plouzané, France