1. Qinghai Academy of Agriculture and Forestry Sciences, Qinghai University, Xining 810016, China
2. Key Laboratory of Agricultural Integrated Pest Management, Xining 810016, China
3. Key Laboratory of Qinghai Tibetan Plateau Biotechnology, Ministry of Education, Qinghai University, Xining 810016, China
4. State Key Laboratory of Plateau Ecology and Agriculture, Qinghai University, Xining 810016, China