1. International Joint Laboratory of Forest Symbiology, College of Forestry, Fujian Agriculture and Forestry University, Fuzhou 350002, China
2. Graduate School of Agricultural and Life Sciences, The University of Tokyo, 1-1-1 Yayoi, Bunkyo-ku, Tokyo 113-8657, Japan
3. College of Forestry, Shandong Agricultural University, No. 61 Daizong Street, Taian 271018, China
4. Asian Research Center for Bioresource and Environmental Sciences, Graduate School of Agricultural and Life Sciences, The University of Tokyo, 1-1-1 Midori-cho, Tokyo 188-0002, Japan
5. Jiangsu Key Laboratory for the Research and Utilization of Plant Resources, Institute of Botany, Jiangsu Province and Chinese Academy of Sciences (Nanjing Botanical Garden Mem. Sun Yat-Sen), No. 1 Qianhuhoucun, Zhongshanmen, Xuanwu District, Nanjing 210014, China