1. College of Life Sciences, Beijing Normal University, Beijing 100875, China
2. State Key Laboratory of Vegetable Biobreeding, Institute of Vegetables and Flowers, Chinese Academy of Agricultural Sciences, Beijing 100081, China
3. College of Plant Protection, Hainan University, Haikou 570228, China
4. Ministry of Education Key Laboratory for Biodiversity Science and Ecological Engineering, Beijing 100875, China