1. School of Marine Science and Engineering, Qingdao Agricultural University, Qingdao 266109, China
2. CAS and Shandong Province Key Laboratory of Experimental Marine Biology, Center for Ocean Mega-Science, Institute of Oceanology, Chinese Academy of Sciences, Qingdao 266071, China
3. Laboratory for Marine Biology and Biotechnology, Pilot National Laboratory for Marine Science and Technology (Qingdao), Qingdao 266237, China
4. University of Chinese Academy of Sciences, Beijing 10049, China