1. Central Region Agricultural Research Center, National Agriculture and Food Research Organization, 2-1-18 Kannondai, Tsukuba 305-8666, Japan
2. Strategic Planning Headquarters, National Agriculture and Food Research Organization, 3-1-3 Kannondai, Tsukuba 305-8604, Japan
3. Centre for Crop and Environmental Science, Harper Adams University, Newport TF10 8NB, UK