1. Department of Geography, University of Victoria, Victoria, BC V8P 5C2, Canada
2. Bamfield Marine Science Center, Bamfield, BC V0R 1B0, Canada
3. Oceans Institute & School of Biological Sciences, University of Western Australia, Perth, WA 6009, Australia
4. Department of Biology, University of British Columbia Okanagan, Kelowna, BC V1V 1V7, Canada