1. Faculty of Agriculture and Life Science, Hirosaki University, Aomori 036-8561, Japan
2. The Shirakami Research Center for Environmental Sciences, Faculty of Agriculture and Life Science, Hirosaki University, Aomori 036-8561, Japan
3. United Graduate School of Agricultural Sciences, Iwate University, Morioka 020-8550, Japan
4. Faculty of Education, Hirosaki University, Aomori 036-8560, Japan
5. Happo-Shirakami Geopark Promotion Council, Akita 018-2502, Japan