1. Department of Range and Watershed Management, Faculty of Natural Resources and Environment, Ferdowsi University of Mashhad, Mashhad 91779-48974, Iran
2. Department of Range Management, Faculty of Natural Resources and Marine Sciences, Tarbiat Modares University, Noor 14117-13116, Iran
3. Organization for Programs on Environmental Sciences, Faculty of Arts & Sciences, University of Tokyo, Tokyo 113-0033, Japan