1. Laboratory of Ichthyology, School of Biology, Aristotle University of Thessaloniki, University Campus, P.O. Box 134, 54124 Thessaloniki, Greece
2. Natural History Museum of Crete, University of Crete, 71305 Irakleion, Greece
3. Independent Researcher, 73134 Chania, Greece
4. Department of Biological Sciences, University of Alberta, Edmonton, AB T6G 1Z2, Canada