1. Department of Reproductive and Behavioral Sciences, Saint Louis Zoo, St. Louis, MO 63110, USA
2. Center for Species Survival, Smithsonian National Zoo and Conservation Biology Institute, Front Royal, VA 22630, USA
3. North of England Zoological Society, Chester Zoo, Upton by Chester CH2 1LH, UK
4. Smithsonian Global Health Program, Smithsonian National Zoo and Conservation Biology Institute, Washington, DC 20008, USA
5. National Museum of Natural History, Smithsonian Institution, Washington, DC 20560, USA
6. Department of Epidemiology of Microbial Disease, Yale School of Public Health, New Haven, CT 06520, USA
7. Veterinary Initiative for Endangered Wildlife, Bozeman, MT 59715, USA
8. Department of Anthropology, The Ohio State University, Columbus, OH 43210, USA
9. College of Public Health, The Ohio State University, Columbus, OH 43210, USA
10. Department of Clinical Sciences, Colorado State University, Fort Collins, CO 80523, USA
11. Wildlife Health Sciences, Smithsonian National Zoo and Conservation Biology Institute, Washington, DC 20008, USA