1. The Laboratory of Forestry Genetics, Central South University of Forestry & Technology, Changsha 410004, China
2. National Engineering Laboratory for Applied Technology of Forestry & Ecology in South China, Changsha 410004, China
3. Faculty of Life Science and Technology, Central South University of Forestry & Technology, Changsha 410004, China
4. Lutou National Station for Scientific Observation and Research of Forest Ecosystem, Yueyang 414000, China
5. Key Laboratory of Subtropical Forest Ecology of Hunan Province, Changsha 410004, China