1. Yunnan Key Laboratory for Integrative Conservation of Plant Species with Extremely Small Populations, Kunming Institute of Botany, Chinese Academy of Sciences, Kunming 650201, China
2. Yunnan International Joint Laboratory for Biodiversity of Central Asia, Kunming Institute of Botany, Chinese Academy of Sciences, Kunming 650201, China
3. Department of Biology, Namangan State University, Namangan 716019, Uzbekistan
4. CAS Key Laboratory for Plant Diversity and Biogeography of East Asia, Kunming Institute of Botany, Chinese Academy of Sciences, Kunming 650201, China
5. University of Chinese Academy of Sciences, Beijing 100864, China
6. Institute of Botany, Academy of Sciences, Tashkent 100125, Uzbekistan