1. Alaska Department of Fish and Game, Threatened, Endangered and Diversity Program, Anchorage, AK 99518, USA
2. School of Natural Resources, University of Nebraska-Lincoln, Lincoln, NE 68583, USA
3. Nebraska State Museum, University of Nebraska-Lincoln, Lincoln, NE 68588, USA
4. U.S. Fish and Wildlife Service, Division of Migratory Birds, Anchorage, AK 99503, USA
5. Environment and Climate Change Canada, Canadian Wildlife Service, Toronto, ON M3H 5T4, Canada
6. U.S. Fish and Wildlife Service, Kanuti National Wildlife Refuge, Fairbanks, AK 99701, USA
7. U.S. Geological Survey, Alaska Science Center, Anchorage, AK 99508, USA
8. Biology Department, Trent University, Peterborough, ON K9L 0G2, Canada
9. U.S. Geological Survey, Nebraska Cooperative Fish and Wildlife Research Unit, School of Natural Resources, University of Nebraska-Lincoln, Lincoln, NE 68583, USA