1. Department of Applied Biology, College of Sciences, University of Sharjah, Sharjah P.O. Box 27272, United Arab Emirates
2. National Research Centre, Genetics & Cytology Department, Cairo 12622, Egypt
3. Tobacco Research Institute, Chinese Academy of Agricultural Sciences, Qingdao 266101, China
4. SINO_PAK Joint Research Laboratory, Institute of Plant Breeding and Biotechnology, MNS University of Agriculture, Multan 60000, Pakistan