1. School of Life Science, Liaocheng University, Liaocheng 252059, China
2. National Engineering Research Center for Vegetables, Beijing 100097, China
3. Beijing Key Laboratory of Vegetable Germplasms Improvement, Beijing 100097, China
4. Key Laboratory of Biology and Genetics Improvement of Horticultural Crops (North China), Beijing 100097, China
5. Beijing Vegetable Research Center (BVRC), Beijing Academy of Agriculture and Forestry Science (BAAFS), Beijing 100097, China