1. College of Life Science and Technology, Huazhong Agricultural University, Wuhan 430070, China
2. Key Laboratory of Protection and Utilization of Biological Resources in Tarim Basin, Xinjiang Production and Construction Corps, College of Life Sciences, Tarim University, Alar 843300, China
3. Hubei Provincial Key Laboratory for Protection and Application of Special Plant Germplasm in Wuling Area of China, College of Life Sciences, South-Central University for Nationalities, Wuhan 430074, China