1. State Key Laboratory of Marine Resource Utilization in South China Sea, Hainan University, Haikou 570228, China
2. School of Life Sciences, Hainan University, Haikou 570228, China
3. College of Ecology and Environment, Hainan University, Haikou 570228, China
4. Analysis and Testing Center, Chinese Academy of Tropical Agricultural Sciences, Haikou 571101, China