1. College of Animal Sciences (College of Bee Science), Fujian Agriculture and Forestry University, Fuzhou 350002, China
2. College of Life Sciences, Fujian Agriculture and Forestry University, Fuzhou 350002, China
3. Laboratory of Evolution and Diversity Biology, UMR5174, University Toulouse III Paul Sabatier, CNRS, 31062 Toulouse, France
4. Biological Science Research Center, Southwest University, Chongqing 400715, China
5. Faculty of Science, University of Queensland, Brisbane, QLD 4072, Australia
6. Mudanjiang Branch of Heilongjiang Academy of Agricultural Sciences, Mudanjiang 157041, China
7. Academy of Bee Science, Fujian Agriculture and Forestry University, Fuzhou 350002, China