1. Institute of Coastal Studies, University Campus of Bragança, Federal University of Pará, Belém 66075-110, Pará, Brazil
2. Museu Paraense Emílio Goeldi, Mastozoology, Belém 66077-830, Pará, Brazil
3. Federal Institute of Education, Science and Technology of Pará, Campus Tucuruí, Tucuruí 68455-210, Pará, Brazil
4. Laboratório de Evolução e Genética Animal, Federal University of Amazonas, Manaus 69067-005, Amazonas, Brazil
5. School of Environment and Life Sciences, University of Salford, Salford M5 4WT, UK