1. Laboratoire Ecologie Fonctionnelle et Environnement, Institut National Polytechnique de Toulouse (INPT), Centre National de la Recharche Scientifique (CNRS), Université de Toulouse (UPS), 31326 Toulouse, France
2. Department of Landscape Ecology and Resources Management, Justus Liebig University, 35390 Gießen, Germany
3. Department of Ecoscience—Catchment Science and Environmental Management, Aarhus University, 8000 Aarhus, Denmark
4. Department Computational Landscape Ecology, UFZ—Helmholtz-Centre for Environmental Research, 04318 Leipzig, Germany