1. College of Animal Sciences, Xinjiang Agricultural University, Urumqi 830052, China
2. Xinjiang Key Laboratory for Ecological Adaptation and Evolution of Extreme Environment Biology, College of Life Sciences, Xinjiang Agricultural University, Urumqi 830052, China
3. State Key Laboratory of Biocontrol, School of Ecology, Sun Yat-sen University, Shenzhen 518107, China
4. Comparative Socioecology Group, Max Planck Institute of Animal Behavior, 78467 Konstanz, Germany
5. Milner Centre for Evolution, Department of Biology and Biochemistry, University of Bath, Bath BA1 7AY, UK
6. Department of Evolutionary Zoology and Human Biology, University of Debrecen, H-4032 Debrecen, Hungary