1. School of Ocean, Yantai University, Yantai 264005, China
2. Guangxi Academy of Sciences, Nanning 530007, China
3. Institute of Beibu Gulf Marine Industry, Fangchenggang 538000, China
4. Graduate School of Fisheries and Environmental Sciences, Nagasaki University, Nagasaki 852-8521, Japan
5. East China Sea Fisheries Research Institute, Chinese Academy of Fishery Sciences, Shanghai 200090, China