1. Department of Marine Ecology and Environment, Gangneung-Wonju National University, Gangneung 25457, Republic of Korea
2. School of Earth Science & Environmental Engineering, Gwangju Institute of Science and Technology, Gwangju 61005, Republic of Korea
3. 21 Century Ocean Development Co., Ltd., Gangneung 25459, Republic of Korea
4. East Sea Branch, Korea Fisheries Resources Agency, Pohang 37601, Republic of Korea