1. Center for Food Animal Health, The Ohio State University College of Food, Agriculture and Environmental Sciences, Wooster, OH 44691, USA
2. Veterinary Preventative Medicine, The Ohio State University College of Veterinary Medicine, Columbus, OH 43210, USA
3. Cleveland Metroparks Zoo, Cleveland, OH 44109, USA
4. Ohio Department of Natural Resources Division of Wildlife, Columbus, OH 43299, USA