1. School of Marine Sciences, Sun Yat-sen University, and Southern Marine Science and Engineering Guangdong Laboratory (Zhuhai), Zhuhai 519020, China
2. Guangdong Provincial Key Laboratory of Marine Resources and Coastal Engineering School of Marine Sciences, Sun Yat-sen University, Guangzhou 510275, China
3. Key Laboratory of Science and Technology on Operational Oceanography, South China Sea Institute of Oceanology, Chinese Academy of Sciences, Guangzhou 510301, China
4. School of Ocean Engineering and Technology, Sun Yat-sen University, Zhuhai 519000, China