1. School of Geosciences, University of Edinburgh, Alexander Crum Brown Road, Edinburgh EH9 3FF, UK
2. UK Centre for Ecology & Hydrology (UKCEH), Bush Estate, Penicuik EH26 0QB, UK
3. Scotland’s Rural College (SRUC), Peter Wilson Building, King’s Buildings, West Mains Road, Edinburgh EH9 3JG, UK