1. Faculty of Biology and Biotechnology, Al-Farabi Kazakh National University, 71 Avenue Al-Farabi, Almaty 050040, Kazakhstan
2. Conservation and Genetic Improvement Center, Camel Project UTF/SAU/044/SAU, P.O. Box 761, Al-Kharj 11942, Saudi Arabia
3. UMR-SELMET, CIRAD-ES, Campus International de Baillarguet, CEDEX 5, 34938 Montpellier, France