1. Department of Biology, School of Biological Sciences, Tokai University, Sapporo 0058601, Japan
2. Department of Obstetrics, Gynaecology and Reproduction, Faculty of Veterinary Science, Chulalongkorn University, Bangkok 10330, Thailand
3. Veterinary Clinical Stem Cells and Bioengineering Research Unit, Chulalongkorn University, Bangkok 10330, Thailand
4. Center of Elephant and Wildlife Health, Faculty of Veterinary Medicine, Chiang Mai University, Chiang Mai 50100, Thailand
5. The Royal Initiated Thai Elephant Conservation Center, Lampang 52190, Thailand
6. Faculty of Veterinary Medicine, Kasetsart University, Nakhon Pathom 73140, Thailand
7. Faculty of Veterinary Science, Prince of Songkla University, Songkhla 90110, Thailand
8. Department of Animal Reproduction, Joint Faculty of Veterinary Medicine, Yamaguchi University, Yamaguchi 7538515, Japan
9. Bio-Innovation Research Center, Tokushima University, Tokushima 7793233, Japan