1. George Mason Centre for the Natural Environment, University of Auckland, Auckland 1010, New Zealand
2. The New Zealand Institute for Plant & Food Research Limited, Mt Albert Research Centre, Auckland 1025, New Zealand
3. The New Zealand Institute for Plant & Food Research Limited, Havelock North 4157, New Zealand
4. School of Biological Sciences, University of Auckland, Auckland 1010, New Zealand
5. Department of Primary Industries and Regional Development, Perth, WA 6000, Australia