1. World Organisation for Animal Health Rabies Reference Laboratory, Agricultural Research Council, Onderstepoort Veterinary Institute, Onderstepoort, Pretoria 0110, Gauteng, South Africa
2. Division for Outbreak Preparedness and Response and the Centre for Emerging, Zoonotic and Parasitic Diseases, National Institute of Communicable Diseases, 1 Modderfontein Road, Johannesburg 2192, Gauteng, South Africa
3. Department of Veterinary Tropical Diseases, Faculty of Veterinary Sciences, University of Pretoria, Onderstepoort, Pretoria 0110, Gauteng, South Africa