1. College of Resources and Environment, Tibet Agriculture and Animal Husbandry University, Nyingchi 860000, China
2. Key Laboratory of Forest Ecology in Tibet Plateau, Tibet Agricultural & Animal Husbandry University, Ministry of Education, Nyingchi 860000, China
3. Linzhi National Forest Ecosystem Observation & Research Station of Tibet, Nyingchi 860000, China
4. School of Forestry, Guangxi University, Nanning 530004, China
5. School of Mechanical Engineering, Guangxi University, Nanning 530004, China
6. International Center for Climate and Global Change Research, College of Forestry, Wildlife and Environment, Auburn University, Auburn, AL 36849, USA