1. School of Forest Sciences, Faculty of Science, Forestry and Technology, University of Eastern Finland (UEF), P.O. Box 111, FI-80101 Joensuu, Finland
2. Bioeconomy and Environment, Natural Resources Institute Finland (Luke), Yliopistokatu 6B, FI-80100 Joensuu, Finland
3. Natural Resources, Natural Resources Institute Finland (Luke), Paavo Havaksen tie 3, FI-90570 Oulu, Finland
4. Finnish Wildlife Agency, Kiekkotie 4, FI-70200 Kuopio, Finland