1. Genome, Cell, and Developmental Biology Graduate Program, Department of Biology, Indiana University Bloomington, Bloomington, IN 47405, USA
2. Medical Sciences Program, Indiana University School of Medicine, Bloomington, IN 47405, USA
3. Indiana University Melvin and Bren Simon Comprehensive Cancer Center, Indianapolis, IN 46202, USA
4. Department of Medicine, Indiana University School of Medicine, Indianapolis, IN 46202, USA
5. Department of Medical and Molecular Genetics, Indiana University School of Medicine, Indianapolis, IN 46202, USA