1. Laboratory of Eremology and Combating Desertification (LR16IRA01), Institut des Régions Arides (IRA), University of Gabes, Medenine 4119, Tunisia
2. Laboratory of Biology & Ecophysiology of Plants in Arid Environments, Faculty of Sciences of Sfax, University of Sfax, Sfax 3038, Tunisia
3. Laboratory of Pastoral Ecosystems and Valorization of Spontaneous Plants and Associated Microorganisms (LR16IRA03), Institut des Régions Arides (IRA), University of Gabes, Medenine 4119, Tunisia