1. (DLJQ, BAB) Integrative Ecology Laboratory, Department of Biology, Faculty of Science, Chulalongkorn University, Pathumwan, Thailand & Center of Excellence in Entomology: Bee Biology, Diversity of Insects and Mites, Chulalongkorn University, Pathumwa
2. (RSC) ICIPE, International Centre of Insect Physiology and Ecology, Nairobi 00100, Kenya & Department of Entomology, U. S. National Museum of Natural History, Smithsonian Institution, Washington, DC 20560, USA (e-mail: rcopeland@icipe.org).