1. (TI) Department of Zoology, National Museum of Nature and Science, 4-1-1 Amakubo, Tsukuba, Ibaraki 305-0005, Japan (e-mail: )
2. (YA) Faculty of Social and Cultural Studies, Kyushu University, 744 Motooka, Fukuoka 819-0395, Japan
3. (CYS and DHZ) Institute of Evolutionary Ecology and Conservation Biology, Central South University of Forestry and Technology, Shaoshan South Road, No. 498, Changsha, Hunan 410004, China