1. Department of Integrative Biology and the University Herbarium, University of California, Berkeley, CA 94720, USA
2. Department of Botany, National Museum of Natural History, Smithsonian Institution, Washington, DC 20560, USA
3. College of Liberal Arts and Sciences, Graduate School of Global and Transdisciplinary Studies, Chiba University, Chiba, Japan
4. Duke University Herbarium, Department of Biology, Durham, NC 27708, USA
5. Boyce Thompson Institute and Plant Biology Section, Cornell University, Ithaca, NY 14853, USA
6. Department of Natural and Applied Sciences and Gary A. Fewless Herbarium, University of Wisconsin, Green Bay, WI 54311, USA