1. Dept. of Environmental Sciences, Faculty of Life and Environmental Sciences, Prefectural Univ. of Hiroshima, 562 Nanatsuka-Cho, Shobara City, Hiroshima 727-0023, Japan.
2. Dept. of Environmental Sciences, Faculty of Life and Environmental Sciences, Prefectural Univ. of Hiroshima, 562 Nanatsuka-Cho, Shobara City, Hiroshima 727-0023, Japan (corresponding author).
3. Environmental Research and Management Center, Hiroshima Univ., 1-5-3 Kagamiyama, Higashi-Hiroshima, Hiroshima 739-8513, Japan.