1. College of Life Science, Shandong Univ., Jinan 250100, China; and College of Population, Resources, and Environment, Shandong Normal Univ., Jinan 250014, China (corresponding author).
2. College of Life Science, Shandong Univ., Jinan 250100, China.
3. School of Economics, Nankai Univ., Tianjin 300071, China.
4. College of Population, Resources, and Environment, Shandong Normal Univ., Jinan 250014, China.