1. Key Laboratory of Ecosystem Network Observation and Modeling, Institute of Geographic Sciences and Natural Resources Research, Chinese Academy of Sciences, Beijing 100101, China; and Graduate Univ. of Chinese Academy of Science, Beijing 100049, China.
2. Key Laboratory of Marine Environmental Science and Ecology, Ministry of Education, Ocean Univ. of China, Qingdao 266003, China.
3. Professor, Key Laboratory of Ecosystem Network Observation and Modeling, Institute of Geographic Sciences and Natural Resources Research, Chinese Academy of Sciences, Beijing 100101, China (corresponding author).