1. Institute for Theoretical Biology, Humboldt-Universität zu Berlin, 10115 Berlin, Germany, and Honda Research Institute Europe GmbH, Offenbach-Main, 63073, Germany
2. Institute for Theoretical Biology, Humboldt-Universität zu Berlin, 10115 Berlin, Germany
3. Institute for Theoretical Biology, Humboldt-Universität zu Berlin, 10115 Berlin, Germany, and Institut für Neuroinformatik, Ruhr-Universität Bochum, 44780 Bochum, Germany