1. Computational Brain Project, ICORP, JST, Sora Ku-gun, Kyoto, 619-0288, Japan; ATR Computational Neuroscience Laboratories, Soraku-gun, Kyoto 619-0288, Japan,
2. ATR Computational Neuroscience Laboratories, Soraku-gun, Kyoto 619-0288, Japan; Initial Research Project, OIST, Gushikawa, Okinawa, 904-2234, Japan; and CREST, JST, Soraku-gun, Kyoto 619-0288, Japan,