1. Computational Neurobiology Laboratory, The Salk Institute for Biological Studies, La Jolla, CA, 92037, U.S.A.
2. Institute for Neural Computation, University of California at San Diego, CA, 92093, U.S.A.
3. Qualcomm, San Diego, CA, 92121, U.S.A.
4. Howard Hughes Medical Institute at the Salk Institute, La Jolla, CA, 92037, and Division of Biological Sciences, University of California at San Diego, CA, 92093, U.S.A.