1. Sloan Center for Theoretical Neurobiology, Salk Institute for Biological Studies, La Jolla, CA 92037, U.S.A. and Georgetown Institute for Cognitive and Computational Sciences, Georgetown University Medical Center, Washington, DC 20007, U.S.A.
2. The Howard Hughes Medical Institute, Salk Institute for Biological Studies, La Jolla, CA 92037, U.S.A. and Department of Biology, University of California San Diego, La Jolla, CA 92037, U.S.A.