1. Natural History Museum and Institute, Chiba, 955-2 Aoba-cho, Chuo-ku, 260-8682 Japan
2. Department of Biological Resources, National Chiayi University, Chiayi, 60004, Taiwan, R. O. C.;, Email: scchang@mail.ncyu.edu.tw.
3. Institute of Marine Biology and Center of Excellence for the Oceans, National Taiwan Ocean University, Keelung 202301, Taiwan, R. O. C.