1. Department of Veterinary Medicine, University of Perugia, via San Costanzo 4, 06126 Perugia, Italy
2. Natural History Museum of the University of Florence, Museo ‘La Specola’, via Romana 17, 50125 Florence, Italy
3. National Reference Centre for Animal Leptospirosis, Istituto Zooprofilattico Sperimentale Lombardia ed Emilia-Romagna ‘Bruno Ubertini’, via Bianchi 9, 25124 Brescia, Italy
4. Institute of Animal Pathology, Vetsuisse Faculty, University of Bern, Länggassstrasse 122, 3012 Bern, Switzerland